Interrelationships and Balance in Small-scale UK Ecosystems

               Heron (Tertiary Consumer)
                           ^
                           |
            Stickleback (Secondary Consumer)
                   ^                ^
                   |                |
         Midge Larva                Tadpole (Primary Consumers)
                   ^                ^
                   |                |
                 Algae            Reeds (Producers)

The Process of Nutrient Cycling

In an ancient woodland like Epping Forest, thousands of fallen leaves usually vanish completely by spring, thanks to 700 species of fungi. The nutrient cycle describes how vital minerals move from the physical environment into living organisms and back again. The Gersmehl model sketches out three main stores (boxes) and transfers (arrows) of nutrients:

               [Precipitation]
                      |
                      v
 +---------+      (Fallout)      +---------+
 | BIOMASS | ------------------> | LITTER  | ---> [Runoff]
 +---------+                     +---------+
      ^                               |
      |                               | (Decomposition)
      | (Plant Uptake)                v
      |                          +---------+
      +------------------------- |  SOIL   | ---> [Leaching]
                                 +---------+
                                      ^
                                      |
                                 [Weathering]

• Biomass: All living organisms. This is the largest store in Epping Forest due to the high density of trees.
• Litter: The topmost layer of dead organic matter. This store peaks in autumn during leaf fall.
• Soil: Nutrients held in the earth, which earthworms help mix.

Nutrients move between these stores via transfers. When leaves fall (fallout), nutrients move from biomass to litter. Decomposers then break down the litter into the soil. Finally, plant roots absorb these nutrients from the soil for new growth (plant uptake). The cycle also has inputs, like precipitation adding nutrients to litter, and outputs, like leaching where rainwater washes nutrients deep out of the soil.

The Impact of Environmental Change

Every time a farmer sprays excessive fertiliser near a waterway, it can trigger a deadly chain reaction underwater. Ecosystems naturally exist in a state of balance or equilibrium, but a change to one component causes a "knock-on effect" or "ripple effect" throughout the entire system.

Changes can be driven by abiotic factors, such as extreme weather. For example, during a drought:

1. Water levels in a pond drop, causing margin plants to dry out and die.
2. The water becomes stagnant and oxygen levels deplete.
3. Starved of shelter and oxygen, insect and fish populations decline.
4. Tertiary consumers, like herons, lose their food source and are forced to migrate.

Changes can also be driven by human actions or biotic factors. If agricultural fertiliser runs off into a pond, it causes an algal bloom. The algae block sunlight, killing bottom-dwelling plants. Decomposers then multiply to break down the dead plants, using up all the oxygen and causing the pond to become anoxic (oxygen-starved), resulting in mass fish death. Alternatively, a biotic change like Ash Dieback disease kills ash trees, directly removing habitats for insects and ultimately starving top predators like owls.